Simon A. Levin received his B.A. from Johns Hopkins University and his Ph.D. in mathematics from the University of Maryland. At Cornell University 1965-1992 , he was Chair of the Section of Ecology and Systematics, and then Director of the Ecosystems Research Center, the Center for Environmental Research and the Program on Theoretical and Computational Biology, as well as Charles A. Alexander Professor of Biological Sciences (1985-1992). Since 1992, he has been at Princeton University, where he is currently George M. Moffett Professor of Biology and Director of the Center for BioComplexity. He retains an Adjunct Professorship at Cornell. His research interests are in understanding how macroscopic patterns and processes are maintained at the level of ecosystems and the biosphere, in terms of ecological and evolutionary mechanisms that operate primarily at the level of organisms; in infectious diseases; and in the interface between basic and applied ecology. Levin is a Fellow of the American Academy of Arts and Sciences and the American Association for the Advancement of Science as well as a Member of the National Academy of Sciences and the American Philosophical Society. He Chairs the Governing Council for IIASA, and co-chairs the Science Board of the Santa Fe Institute. Levin is a former President of the Ecological Society of America and the Society for Mathematical Biology, and a past Chair of the Board of the Beijer Institute of Ecological Economics. Among other awards, he won the MacArthur Award (1988) and the Distinguished Service Citation (1998) of the Ecological Society of America, and the Okubo Award of the Society for Mathematical Biology and the Japanese Society for Theoretical Biology. Most recently, he was honored with the Dr. A.H. Heineken Prize for Environmental Sciences by the Royal Netherlands Academy of Arts and Sciences (2004), the Kyoto Prize in Basic Sciences (2005) by the Inamori Foundation, and the Distinguished Scientist Award of the American Institute of Biological Sciences. Levin has mentored more than 100 graduate students and postdoctoral fellows.